Systems with two or more resources
One of the more basic extensions of theory regarding coexistence in varying environments involves systems with two or more resources (point (iv) in the list in Section 9.5.3). There has been a limited amount of work on coexistence in systems with two resources in which cycles are caused purely by consumer-resource interactions (Egas et al.
2004; Abrams 2006a, b, 2007b). Most of this has been discussed in the context of the coevolution of specialist and generalist species. In spite of having identical ranges of evolutionarily possible functional response parameters, coexistence of three species was possible over a wide range of relative mortality rates at the evolutionary equilibrium. Some of the best possibilities for coexistence arose from features that were not possible in the single-resource model studied here; i.e. adaptive adjustment of relative consumption rates by the consumers (Abrams 2006b), and competitive interactions between biotic resources (Abrams 2007b).The shared-exclusive version of the 2-consumer-3-resource models considered in previous chapters has not, to my knowledge been extended to consider the impacts of sustained variability affecting either the shared or exclusive resources. Analysing this system should provide another perspective on the range of impacts of sustained variability on the strength of interspecific competition.
9.7