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Anything that influences energy gain by photosynthesis has the potential to affect the survival, growth, and reproduction of the organism.

As we have just seen, rates of photosynthesis are influenced by environmental conditions,

particularly temperature and water availability. In addition, an apparent biochemical inefficiency in the initial step of the Calvin cycle limits energy gain by photosynthetic organisms.

In this section, we will examine some evolutionary responses to these environmental constraints on photosynthesis. We will describe two specialized photosynthetic pathways, the C4 pathway and crassulacean acid metabolism (CAM), that make photosynthesis more efficient under particular potentially stressful environmental conditions. Plants that lack these specialized pathways use the C3 photosynthetic pathway. The C3 and C4 photosynthetic pathways take their names from the number of carbon atoms in their first stable chemical products. First, we'll examine photorespiration, a process that operates in opposition to the Calvin cycle and lowers its efficiency.

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Source: Bowman W., Hacker S.. Ecology. 6th ed. — Oxford University Press,2023. — 744 p.. 2023

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